Dogs have scent glands in their paws. When they kick up grass after they pee or defecate, for instance, they are imparting their scent to the scattered grass. Paw scent glands provide another marking mechanism for them. Presumably, their evolutionary relatives, such as coyotes and wolves, and other mammals, also have paw scent glands.
When running, the animal's feet lay down a trail of scent, marking the route and therefore making a sensible path for members of the pack. The path could also mark a territorial boundary. Scent routes can be, therefore, both boundaries in the landscape and paths through the landscape. It is possible (I am only suggesting, I have no factual evidence) that paw scents differ according to the structure laid down--whether boundary or path.
Some environmental structures set up animal marking patterns. For instance, upthrust geological strata often weather at different rates, leaving one stratum less eroded than neighboring strata. The exposed, less eroded stratus easily becomes a structure along which canines could run unobstructedly, which invites such running. Deposit of a scent trail marks the geological prominence as a boundary and/or path, bring a coincidence between geology and canine landscape.
Similarly, the tops of ridges provide gutters for rain runoff. Over years of runoff, there is less vegetation on the ridgetop, because rain washes away seeds and seedlings. Persons who have hiked in the wilderness have undoubtedly noticed these natural pathways. The ridgetop "paths" invite animals to take them as routes. They provide good observation on either side for predators. They provide rapid escape routes.
Geological structures thereby become boundaries and pathways for animals. When man comes along, animal behavior compels humans to acknowledge, unconsciously, the animal topography. If hunting an animal that has created a bounded topography in the landscape, humans will act with reference to that topography. When humans win ecological dominance in the local environment, they will have unconsciously adopted much of the animal topography of the chief animals on which their society is based.